I think this is more of a replacement for the 200-600mm. Add a TC 1.4 and you get a 180-600mm f6.3 that would probably be just as good, if not better than the 200-600mm f6.3. Take the TC off, and you have a very good 100-400mm lens. If you happen to use both the 200-600mm and 100-400mm lenses, your overall bag can end up even lighter. Add a TC 2x, and you will probably get a good 200-800mm f9 lens. This could be one lens that covers most wildlife needs.
Weight aside, I could also see people pairing this with the 400-800mm. 100-400mm f4.5 for low light and close subjects, and 400-800mm f8 for max reach.
Overall, I think Sony is going more after Nikon's 200-400mm f4 ish lenses. Those lenses are about 8lbs, longer than the 200-600mm, and 10k+. If a 1/3 less of a stop gets us 100mm more on the wide end, a shorter lens, decent weight savings, and less than 1/2 the cost, this lens should be a real winner.