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The 2nd reply that I have from Nikon UK.
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Subject: D5 and D500 - Dynamic AF not working correctly | Case #: 01867793
Response from John M
Further to your recent inquiry I have some additional information which I hope explains what you are experiencing.
The AF engine in the D5 and D500 has changed significantly to the Multi-CAM 20K AF sensor module, a key difference with this is that besides the significant increase in focusing points it also has its own dedicated processor. Due to all of these different components there are differences compared to previous cameras.
1. What custom option is set for a3 in the camera menu, with the D5 and D500 you get additional control depending on the subject matter. The higher density of AF points can be a benefit or depending on the subject matter reducing the number of dynamic AF points can be good too. See the guide below for further information on how this might be helpful.
AF lock-on can be set with two parameters:
“Subject motion" to set characteristics of the subject’s movement
"Blocked shot AF response" to set sensitivity of response to any object (other than the subject) moving across the frame
Simple setting terms ("Erratic/steady," “Quick/Delayed") are adopted for more intuitive setting
2. The main AF point in Dynamic AF can be moved around and this is recorded where ever it was at the time of exposure after recomposing.
3. Different components are used and listed in the User Manual and its normal for performance to vary between two products in this scenario, not everything is listed in the user manual, but the users guides from our download centre can expand further with this along with asking us directly.
I hope this reply clarifies things and if you have any other queries for Nikon in the future please contact us again.
Nikon Customer Support